Palaearctic species is mainly distributed in the southern half of
Europe (Schaffner et al. 2001). In Eastern Europe the species is
spread from the southern Ukraine and the Volga delta through middle
and southwest Asia to Iran and Pakistan (Becker et al. 2003). It
is rarely found in Central Europe (Aspöck 1969, Schaffner et
stages develop in shallow herbaceous marshes, temporary pools, irrigation
ditches or the watersides of lakes, overgrown with duckweed or straight
vegetation and rich in organic matter; they inhabit fresh or slightly
brackish waters, which may be sunlit or slightly shaded (Schaffner
et al. 2001, Becker et al. 2003).
can be found from May to the beginning of October, reaching highest
densities in August (Becker et al. 2003). In southern countries
larvae may be present throughout the year (Schaffner et al. 2001).
In the delta of the river Po larvae, pupae and imagoes have been
collected at same time in late spring. Adults are most abundant
in late summer (Becker et al. 2003). In Austria females have been
encountered in low abundance in September (Aspöck et al. 2001).
According to Schaffner et al. (2001) and Becker et al. (2003) it
is probably the females, which overwinter in the colder regions.
do not bite humans or other mammals, and seem to be autogenous;
the species has no known medical importance (Schaffner et al. 2001).